Dec-2022: Completed a comprehensive review and revision of Torreya, which is a fascinating genus; nearly every species has at least one remarkable story. Posted an updated piece on Future Forests of Washington. Revised many species of Juniperus, particularly in the J. oxycedrus and J. virginiana species complexes. I did this in part to fulfill a promise to Bob Adams, who passed away earlier this year. Most recently, I prepared a very comprehensive page on Tetraclinis, which has one of the oldest and most complex records of human use.
Aug-2022: Eric Van Buskirk has provided a Topics article on the Japanese practice of Forest Bathing and some of its unexpected health benefits.
Apr-2022: I gave a presentation on the Future Forests of Washington, now on Youtube (here's the presentation file). The bottom line is, climate change is bad news for forests, to the point where nearly every tree species is going to need human help if it is going to survive the next century. Washington is a microcosm of the global ecological disaster. You should watch this, because this is based on peer-reviewed science and it shows that the popular media still doesn't understand how severe this problem is, because they report the little pieces of bad news but not the big picture. Here's the big picture.
Dec-2021: New page on the history of conifer botany. Zillions of minor changes as I try to update the conservation status and provide citations to original descriptions of all taxa. EXTENSIVE revision of the group of species formerly known as Pinus ponderosa. Genetic and morphological work found the group to be paraphyletic and it has been subdivided into 3 subspecies of ponderosa, plus 2 "new" species, Pinus brachyptera and Pinus scopulorum (which, BTW, is Latin for "pine of the Rockies"). Made a new ponderosa map.
Oct-2021: I was interviewed on the popular plant podcast In Defense of Plants, talking about old trees in general and especially about some exciting new genetic research on longevity, recently done in Welwitschia. Give it a listen. Also, revised Retrophyllum and its Malesian species. Other new updates include a page on conifers and beer, extensive revisions to Pinus monticola, and lots of minor updates and new photos. More to come.
Some Other Links
Besides the Topics page, here are a couple of other links to get you started:
My curriculum vita (PDF file)
Welcome to the Gymnosperm Database, the web's premier source of information on conifers and their allies. Since we went online on 1997, the Database has attracted worldwide attention as a readily accessible, scientifically accurate source of information on the classification, description, ecology and uses of this culturally and ecologically important group of plants. This home page gives some hints about how to navigate the Database, as well as providing background information for the curious.
There are three major sections of the site:
| Species | Descriptions of all the species and higher-rank taxa of gymnosperms. For each taxon I provide information on classification, description, ecology, ethnobotany, and various other topics of interest. |
|---|---|
| Topics | Things that are not strictly tied to a particular species. For instance, pages on the gymnosperms of Australia (and other regions), the largest and oldest trees in the world, and accounts of some of my field excursions. |
| Bookstore | Most of the books sold here are linked through Amazon, and if you buy them through this site, the price is the same to you but some of the proceeds come to me, to help support the Database. |

Shade foliage of Abies procera.
The Gymnosperm Database was established as an online entity in the summer of 1997 and has since grown steadily, getting its own URL (Conifers.org) in the summer of 1999. The Database provides information for all species and higher-ranked taxa of the gymnosperms, i.e., conifers, cycads, ginkgo, and the gnetophytes. You enter the taxonomic tree at the Family level and then navigate to the Genus, Species or sometimes Variety levels. At each level, information on the taxon is provided, along with bibliographic citations that will take you to more detailed information about the taxon.
Description of Database Fields.
If you have a specific question, use the search box (top of page at right). Otherwise, I suggest you start with the Topics page, or browse the family, genus, and species accounts. The site covers conifers, ginkgo and gnetophytes (Ephedra, Gnetum, and Welwitschia) in detail. Although they are gymnosperms, in recent years I have neglected the cycads because an excellent site, the World List of Cycads, exists to provide the same sort of information that I would otherwise have to collect.
For all nations I use the common English name (e.g., "Spain" instead of "España"), but for local names I try to use the local language if it can be done in the Roman alphabet (e.g. Nuevo León); otherwise I use a general Romanized name such as is found on Google Maps, sometimes also showing the name in the relevant local language. I try to name every nation that each species is native to. For species native to Australia, Canada, China, India, Mexico, and the United States (and sometimes other countries), I try to name every state or province that each species is native to. Abbreviations are not used. This information will help you in geographic searches. For instance, to find all the gymnosperms native to Iowa, simply type in the search term "Iowa."
The Database is not a "real" database. It simply consists of a large collection of HTML files (W3C compliant HTML 4.01, put together with PHP). The benefit of this is that I spend more time writing about gymnosperms and less time programming. The drawback is that I get a lot of spam from people telling me I have to pay them to rewrite all 1500 of my pages in XTML.
Links to citations pop up an a small window, like this. Links to photos also generally pop up in a window, though a few take you offsite. Onsite links take you directly to the linked page, while offsite links open in a new browser window (or a new tab).
One consequence of the longevity of the database is that the graphics quality varies quite a bit. When I started out, everybody was working on a 640×480 pixel screen and images larger than 20 KB took forever to download. So, some of the old images on the site are quite small. Currently the site is laid out to fit a 1024-pixel width (which works well for browsers, smartphones, and hardcopy printing) and some of the pictures are over 100 KB, although pictures larger than 10 KB are only shown in pop-up windows. I entered digital photography in 2001 and the hardware has steadily improved since that time.
People sometimes write and say, "Who writes the database?" The answer is, I do. I have assembled most of the database and done essentially all of the computer-based work: typing, scanning images, writing code, etc. Any assistance is welcome! To date I have received helpful advice from a variety of recognized taxonomic authorities, as well as information on individual taxa from persons scattered across the globe. Many visitors have sent photos or even fine art drawings of gymnosperms. In fact, I have to say that running this website has introduced me to new friends, provided opportunities to travel to memorable places, and provided opportunities to share both my writing and photography. The correspondence related to this site provides one of the greatest rewards of maintaining this site. However, if you would like to increase those rewards, you could buy some books (from Amazon.com) via my gateway/bookstore located HERE.
Last Modified 2022-12-29