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photograph

Original illustration (Kirk 1889).

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Tree ca. 100 cm diameter in Puketi Forest [C.J. Earle, 2003.03].

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Bark on a tree ca. 40 cm diameter in Tongariro National Park [C.J. Earle, 2003.03].

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Foliage of an adult plant on Hauhungatahi, Tongariro National Park; longest leaves are 2 cm long [C.J. Earle, 2003.03].

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Foliage of a juvenile plant in Puketi Forest; longest leaves are 7 cm long [C.J. Earle, 2003.03].

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Seeds (Webb and Simpson 2001).

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Distribution map (Metcalf 2002).

 

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Conservation status 2010: protocol 2.3, needs updating

Podocarpus laetus

Hooibr. ex Endl. 1847

Common names

Montane totara (Salmon 1996), thin-bark totara, Hall's totara (Metcalf 2002), tōtara kōtukutuku [Maori] (Simpson 2017). See also the totara, P. totara.

Taxonomic notes

Synonymy:

Until 2015, this species was known variously as Podocarpus cunninghamii or P. hallii, and virtually all of the scientific literature to that time uses one of these two names. It was finally discovered that a specimen described in 1847 as having been collected in Australia was actually a representative of this taxon and the name assigned to that specimen, P. laetus, is now the valid name by reason of priority (Molloy 2015).

Description

Tree to 20 m tall and 125 cm dbh. Bark thin, papery. Branchlets slender, somewhat drooping on juvenile trees. Leaves brown-green; juvenile foliage 3-7 cm long by 3-5 mm wide, linear-lanceolate; adult foliage smaller, 1.5-3 cm long by 3-4 mm wide, narrow-linear to linear-lanceolate, acute to acuminate, coriaceous, pungent, the midvein usually evident. Pollen cones are 1-2.5 cm long, solitary or up to 5 on a common peduncle, with 4 scales, apiculus obtuse; falling soon after pollen is shed. Ovules solitary or paired; receptacles usually red, swollen and succulent. Seed green, 3-5 mm long, narrow-ovoid, pointed, nutlike (Allan 1961, Metcalf 2002, pers. obs. 2003).

Metcalf (2002) asserts that it is most easily distinguished from P. totara, with which it often grows, by its thin and rather papery bark as opposed to the tough, thick and fibrous bark of P. totara. Matsui et al. (2004) generally agree with that, but present evidence of exceptions; as usual, a confident identification is based on a variety of characters.

Distribution and Ecology

New Zealand: N Island, S Island and Stewart Island (Farjon 1998). Based on data from 1456 collection localities, its climate preferences include a mean annual temperature of 9.2°C, with an average minimum in the coldest month of -0.8°C, and a mean annual precipitation of 2539 mm (Biffin et al. 2011, Table S5). Hardy to Zone 8 (cold hardiness limit between -12.1°C and -6.7°C) (Bannister and Neuner 2001).

Both this and P. totara are tall, slow-growing forest trees that often grow together in lowland forests from sea level to 600 m, with P. totara giving way to P. hallii above 480 m (Salmon 1996). I have seen it growing in what appear to be two distinct ecological niches in the North Island. The first role is that of montane totara, where it grows as a forest dominant on well drained soils at elevations above 480 m. The second role is that of a codominant tree tolerant of wet soils, in which capacity it grows on very wet substrates at low elevations throughout much of Northland. Examples include trees growing on wet floodplain soils in Waipoua and Trounson Kauri Forests, and trees growing very near mangroves around Auckland.

Big tree

A tree 30 m tall and 265 cm dbh, the 'Motu totara,' is reported from Dean Forest, Southland (Burstall and Sale 1984).

Oldest

Smale and Smale (2003), studying forest dynamics and tree growth rates, report ages of up to 496 years for trees growing in native forest remnants at Waihaha in the central North Island. Ages in this study were determined by simple ring-counts of increment cores taken at breast height.

Dendrochronology

This species has been used in at least one climate change study (Burrows and Greenland 1979), and was probably the first New Zealand species to see any dendrochronological use, in an investigation of Maori archeology (Batley 1956).

Ethnobotany

The bark "was formerly used by southern Maori to encase the kelp bags used for storing muttonbirds. These containers are known as pohatiti" (Metcalf 2002).

Observations

In Tongariro National Park it grows on the track up Hauhungatahi, a subsidiary cone west of Mount Ruapehu. To reach this stand, drive 5.5 km south from the highway junction in National Park to the tiny hamlet of Erua and park near the railroad tracks. Follow the tracks about 250 m south to where a small, weathered sign says 'Track' and follow it. The track heads almost continuously uphill, starting in a flax thicket and then, in the forest, passing progressively through elevation zones dominated by rimu (Dacrydium cupressinum), matai (Prumnopitys taxifolia), montane totara (Podocarpus hallii), kaikawaka (Libocedrus bidwillii), yellow pine (Halocarpus biformis), mountain toatoa (Phyllocladus alpinus), bog pine (Halocarpus bidwillii) and pygmy pine (Lepidothamnus laxifolius). This species can also be seen growing along an elevational sequence along the Tongariro Crossing, above the Ketetahi trailhead. As noted above, it also grows in very wet soils in Northland; good examples can be seen in Trounson Kauri Forest, along the ridge walk in Puketi Forest, and along the walk to Te Matua Ngahere (the biggest kauri) in Waipoua Forest.

Remarks

Citations

Batley, R.A.L. 1956. Some practical aspects of dendrochronology in New Zealand. Journal of the Polynesian Society 65(3): 232-244 (as Pinus hallii).

Molloy, Brian. 2015. The correct name for the New Zealand endemic conifer Hall’s totara (Araucariales: Podocarpaceae). Phytotaxa 220(2):101-116.

Simpson, Philip. 2017. Totara: A Natural and Cultural History. Auckland University Press, 300 pp.

See also

Matsui, T., J.B. Wilson, and C.J. West. 2004. Can Podocarpus totara and P. hallii be distinguished by bark thickness?: A study on the southern coast of Southland/Otago, New Zealand. New Zealand Journal of Botany 42:313-320. http://www.royalsociety.org.nz/Site/publish/Journals/nzjb/2004/020.aspx, accessed 2009.03.23, now defunct).

Smale, M.C. and P.N. Smale. 2003. Dynamics of upland conifer/broadleaved forest at Waihaha, central North Island, New Zealand. Journal of the Royal Society of New Zealand 33(2):509-528. http://www.royalsociety.org.nz/Site/publish/Journals/jrsnz/2003/024.aspx, accessed 2009.03.23, now defunct).

Wardle, P. 2001. Distribution of native forest in the upper Clutha district, Otago, New Zealand. New Zealand Journal of Botany 39:435-446. http://www.royalsociety.org.nz/Site/publish/Journals/nzjb/2001/34.aspx, accessed 2009.03.22, now defunct).

Wardle, P. 2001. Holocene forest fires in the upper Clutha district, Otago, New Zealand. New Zealand Journal of Botany 39:523-542. http://www.royalsociety.org.nz/Site/publish/Journals/nzjb/2001/42.aspx, accessed 2009.03.22, now defunct).

The New Zealand Plant Conservation Network, accessed 2010.11.22.

Gymnosperms of New Zealand.

Last Modified 2017-06-01